¹Neuroscience Program, University of Michigan, Ann Arbor, ²Cellular and Molecular Biology Program, University of Michigan, Ann Arbor, ³Department of Mechanical Engineering, University of Michigan, Ann Arbor, ⁴Department of Biomedical Engineering, University of Michigan, Ann Arbor, ⁵Department of Biophysics, University of Michigan, Ann Arbor

¹The Niels Bohr Institute, University of Copenhagen, ²Department of Plant and Environmental Sciences, University of Copenhagen

¹Department of Applied Chemistry, School of Applied Science, National Defense Academy of Japan, ²Department of Applied Physics, School of Applied Science, National Defense Academy of Japan, ³Department of Materials Science and Technology, Faculty of Engineering, Niigata University

¹Centre for Mechanochemical Cell Biology and Division of Biomedical Sciences, Warwick Medical School, University of Warwick, ²National Centre for Biological Sciences, Tata Institute of Fundamental Research